Modifiers
Grey, Roan, Pinto, and Appaloosa
![]() |
![]() |
![]() |
| And here is Unbridled's Song, another grey who has not developed flea bites. There has been some speculation that fleabites were the result of either heterozygous or homozygous grey, but there doesn't seem to be much evidence to support it. Both Holy Bull and Unbridled's Song are heterozygous greys. |
![]() |
![]() |
![]() |
![]() |
![]() Some bay and chestnut based greys go through a phase called rose grey, like the horse above. Some rose greys dapple out and some, especially Arabians, tend to lighten up quickly without really dappling while still retaining a rosey hue. |
|
| Roan: The roan gene acts by interspersing white hairs throughout a horse's coat, but the head, legs, mane, and tail remain "dark." Roan can act on any of the base colors as well as on any dilute or patterned color. For example, you could have a red roan (chestnut + roan), a strawberry roan (bay + roan), a blue roan (black + roan), or even a palomino roan (chestnut + cream + roan). Roan is often confused with grey, but it should be noted that roan horses look the same throughout their lives while grey horses fade out to nearly white (as seen above). |
|
Chestnut Roan: Chestnut + Roan |
Bay Roan: Bay + Roan |
Black Roan: Black + Roan |
|
A bay roan foal. Notice how his head and points are dark right from birth. |
A palomino roan (chestnut + cream + roan). Even with 2 different dilution genes at work, this horse's head and knees retain traces of the dark palomino color. |
|
Pinto Patterns: Pinto is a term that refers to several different patterns of white. The patterns fall into two catagories, tobianos and overos. The overo catagory includes frame overo, sabino, and splash white. A horse with a combination of tobiano and one or more overo patterns is known as a tovero. The tobiano pattern is dominant, and it is characterized by white crossing the horse's back between the withers and the dock, as well as by areas of color on the head, chest, and flanks. Most tobianos have normal face markings like stars and blazes, and they almost always have four white feet. Frame overos are characterized by having a dark topline between the withers and dock, with blocks of white on their sides and necks. Frame overos usually have large blazes or apron markings.The feet are usually dark, but may be white if combined with the tobiano, sabino, or splash genes. Sabino is a pattern that comes in many forms. In its minimal form, it only manifests itself as small socks and a star or blaze, if that. More extreme sabinos will have white patches, often with roaned edges, creeeping up their sides from the underside of the belly. The most extreme sabinos will look entirely white or almost so, often retaining a few patches of color along their toplines, particularly on the ears or in the mane. Splash white horses look as if they've literally been splashed with white paint from the underside or dipped in white paint. Splash white is an incomplete dominant. Homozygous spashes will have more white than heterozygous splashes. Splash markings tend to have smooth, crisp edges, and most splashes have blue eyes (especially the homozygous ones). All pinto patterns can range from minimal expressions (a nearly solid colored horse) to extreme expression (an all-white or nearly all-white horse). White horses resulting from tovero and sabino patterns are completely viable. White horses that are homozygous for frame overo, however, are known as lethal white overos (LWO). For more information on lethal white, please read my page about white horses. |
Tobiano
|
|
|
|
|
|
![]() This is The Eagle's Gift, a lovely tobiano Paint mare who exhibits cat tracks, the small spots on her shoulder. Cat tracks are often somewhat roany looking and are typical of homozygous tobianos. (Bred and photographed by Spotted Fawn Paints) |
![]() As I mentioned above, nearly all tobianos have 4 white feet. Occassionally, the pattern gets skewed, resulting in tobianos with dark feet like this guy. These skewed patterns are quite unusual and are usually only seen in Miniature Horses and Shetland Ponies. |
Overo
|
|
|
|
|
|
Sabino
|
|
|
|
|
|
Splash White
|
|
|
|
![]() A more extremely marked splash white. Definitely
a homozygous splash.
|
Tovero
|
|
|
|
| Appaloosa: According to recent studies, the appaloosa pattern seems to manifest itself similarly to the pinto genes. Moderately expressed appies will have a small lacy blanket, more boldly marked horses will have larger blankets, and the most boldly patterned horses are entirely covered by the blanket, meaning they are leopard appaloosas. For more information on the ongoing project of mapping the appaloosa genome, please check out The Appaloosa Project. Appaloosa pattered horses have distinct markings besides their spots---they have striped hooves, mottled skin, and white sclera. (This section is under construction while I learn more about Appy genetics.) |
![]() A small blanket. |
![]() A larger blanket. |
![]() A larger blanket, sometimes called semi-leopard. |
|
|
![]() Dudes Bonanza, a varnish roan appaloosa. |
![]() |
![]() |
![]() |